SOLON -- Biologists are trying to figure out what led to the deaths of hundreds of stocked trout in three-acre Shadow Lake in the South Chagrin Reservation of the Cleveland Metroparks.

Metroparks Fisheries biologist Mike Durkalec says oxygen depletion can lead to fish kills when lakes ice over in the winter, but that's doesn't appear to be the cause at Shadow Lake.

He says officials are trying to determine whether the oxygen loss was somehow accelerated by runoff with a "high organic content" but that any comment on the cause of the fish kill would be speculation.

A representative for Waste Management, the company that runs a now-closed landfill near the park, says it's cooperating with investigators.